Defending champion Serena Williams followed sister Venus out of the French Open on a dramatic fourth day.
Serena lost 6-2 6-2 to Spain’s Garbine Muguruza after Venus had been beaten 2-6 6-3 6-4 by Slovakian Anna Schmiedlova at Roland Garros.
It is the first time in her career that Serena has failed to reach the second week of a Grand Slam as the top seed.
It means a projected third-round clash between the Williams sisters will now feature Muguruza and Schmiedlova.
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